MYSMENIDAE Petrunkevitch, 1928

Nombre común en ingles: Minute clasping weavers

Nombre común en castellano: Se desconoce

Familias similares:

Género tipo: Mysmena Simon, 1894

Diagnosis: Very small araneomorph spiders; three tarsal claw; ecribellate; entelegyne; eight eyes; male with mating spine on metatarsus I (sometimes on tibia I as well); cymbium of male palp with lobes or apophyses; female and sometimes males with a sclerotized subdistal ventral spot on anterior femora.

Caracteres descriptivos: Caparazón: usually ligh, with highest point behind eyes, except in Isela and Kilifina where carapace flattened. Esternón: truncated, rarely pointed (Maymena), between coxae IV. Ojos: eight; in two rows; anterior median eyes usually larger than other eyes; lateral eyes contiguous, situated close to anterior median eyes. Quelíceros: usually with tiny denticles scattered between cheliceral teeth. Piezas bucales: labium rebordered. Patas: three claws; male with mating spine on metatarsi I; femur I, and sometimes II of female and sometimes male with sclerotized spot ventro-subdistally; tarsi longer than or equal in lengh to metatarsi; tarsi IV without serrated setae; tarsal organ capsulate. Palpo femenino: not reduced but lacking a claw. Abdomen: soft; spherical to hight than long; usually bearing scattered, long setae; scutum sometimes present. Hileras: anterior lateral spinnerets with reduced bases of piriform gland spigots; posterior lateral spinnerets with triplet; colulus present. Sistema respiratorio: booklungs usually absent. Genitalia: entelegyne; epigyne usually with simple posterior plate or scapus; cymbium of male palp with lobes or apophyses. Tamaño corporal: < 3 mm. Color: varies from yellow-brown to grey, sometimes with greenish tinge, abdomen dark grey with pale markings or white spots.

Estatus taxonómico: After having been considered part of the Theridiidae, Mysmeninae was for some time a subfamily of the Symphytognathidae. Forster & Platnick (1977) accorded the taxon family rank. Coddington & Levi (1991) placed the mysmenids in the Araneoidea and suggested that they are part of the 'higher araneoids', as sister to Symphytognathidae, in a group of families considered to be the sister-group of the Araneidae. Griswold et al. (1998), followed by Coddington et al. (2004). place the Mysmenidae within the so-called Symphytognathoidea, sister to Anapidae + Symphytognathidae. Schütt (2003) corroborates this placement but claims that the mysmenids must be relimited and the Old World genera transferred to the new family Synaphridae, in accordance with Wunderlich (1986). Wunderlich's (2004) opinion that Anapidae should be widened to include the Mysmenidae in only poorly founded by a 'possible' cladogram.

Distribución: mundial.

Estilo de vida: Little is known about the behaviour of mysmenids. Some species are kleptoparasites on the webs of other spiders, whereas others spin sheet webs in low vegetation. Mysmenidae usually inhabit crevices or leaf litter in very humid habitats. Webs are known for a few species and vary from a complex three-dimensional orb with a proliferation of out-of-the-plane radii over a roughly egg-shapped space web to some kind of sheetweb.

Bibliografía:

  • Dippenaar-Schoeman, A.S. & Jocqué, R. 1997. African Spiders: An Identification Manual. Plant Protection Res. Inst. Handbook, no. 9, Pretoria, 392 pp.

  • Forster, R.R. & Platnick, N.I. 1977. A review of the spider family Symphytognathidae (Arachnida, Araneae). Am. Mus. Novit. 2619: 1-29.

  • Gertsch, W.J. 1960. Descriptions of American spiders of the family Symphytognathidae. Am. Mus. Novit. 1981: 1-40.

  • Griswold, C.E., Coddington, J.A., Hormiga, G. & Scharff, N. 1998. Phylogeny of the orb-web building spiders (Araneae, Orbiculariae: Deinopoidea, Araneoidea). Zool. J. Linn. Soc. 123: 1-99.

  • Lopardo, L. & Coddington, J.A. 2005. Mysmenidae. pp. 175-177 in Ubick, D., Paquin, P., Cushing, P.E. & Roth, V. (Eds) 2005. The Spiders of North America. An Identification Manual. American Arachnological Society.

  • Platnick, N.I. & Shadab, M.U. 1978b. A review of the spider genus Mysmenopsis (Araneae, Mysmenidae). Am. Mus. Novit. 2661: 1-22.

  • Schütt, K. 2003. Phylogeny of Symphytognathidae s.l. (Araneae, Araneoidea). Zool. Scripta 32: 129-151.

  • Wunderlich, J. 1986. Spinnenfauna gestern und heute: Fossile Spinnen in Bernstein und ihre heute lebenden Verwandten. Quelle & Meyer, Wiesbaden, 283 pp.

  • Wunderlich, J. 2004. Fossil spiders in amber and copal. Joerg Wunderlich Verlag. Hirschberg-Leutershausen (Germany). Vol. 3 A, B, 1908 pp.

          1 género y 2 especies presentes en la Península Ibérica:

  • Mysmena Simon, 1894

                      - M. gibbosa Snazell, 1986
                      - M. leucoplagiata (Simon, 1879)
 
Agelena labyrinthica © Pedro Cardoso