PIMOIDAE WUNDERLICH, 1986

Nombre común en ingles: Large hammock-web spiders

Nombre común en castellano: Se desconoce

Familias similares:

Género tipo: Pimoa Chamberlin & Ivie, 1943

Diagnosis: Medium to small araneomorph spiders; ecribellate; entelegyne, eight eyes; three tarsal claws; recognised by the presence of a retrolateral cymbial sclerite (piramoid cymbial sclerite), a dorsoectal cymbial process, and cuspules (modified macrosetae) on either the cymbial process or the dorsal surface of the cymbium and a paracymbium attached to the cymbium on the male palp and an epigyne with a scape with the copulatory openings at the distal end.

Caracteres descriptivos: Caparazón: longer than wide elongate conspicuous. Esternón: longer than wide, projecting between coxae IV. Ojos: eight subequal, in two rows; anterior row recurved, posterior row variable (from above); lateral eyes touching; secondary eyes with canoe-shaped tapetum. Quelíceros: large, promargin with two or three, retromargin with 1-4 teeth; usually with stridulating file. Piezas bucales: endites almost retangular, parallel; labium wider than long. Patas: 1423, longer in male than in female; tarsi with three claws; spination variable; usually one trichobothrium on metatarsi; autospasy at patella-tibia junction. Palpo femenino: slender, with numerous spines: tarsal with claw provided with row of minute denticles and basal tooth. Abdomen: oval, slightly longer than wide. Hileras: six, all well developed; colulus fairly large with setae; anterior lateral spinnerets with reduced bases of piriform gland spigots; posterior median spinnerets without aciniform gland spigots; posterior lateral spinnerets with two cylindrical gland spigots (base of peripheral cylindrical spigot slightly larger than base of distal one); triplet absent in males. Sistema respiratorio: two booklungs and tracheal spiracle just in front of spinnerets; tracheae haplotracheate. Genitalia: entelegyne; epigyne large with well developed scpe-like protrusion, consisting of a dorsal and ventral plate, bearing distal copulatory opening; spermathecae spherical; cymbium of male palp with alveolus near prolateral margin, provided with a retrolateral sclerite (in some species fused to paracymbium), a dentate dorsal excrescence and modified macrosetae (most often as denticles) on either cymbial process or dorsal surface of cymbium: paracymbium fused to cymbium (intersegmental in Weintrauboa); embolus long and filiform, continuous with tegulum, paralleled by a long process (absent in Nanoa); tegulum large with membranous conductor and sometimes a median apophysis. Tamaño corporal: 1,4-12 mm. Color: carapace often light brown with darker margins; sternum usually dark; abdomen dark grey with lighter chevron pattern.

Estatus taxonómico: Although a small family, it is difficult to provide a concise definition of it. The Pimoidae are considered the sister-group of the Linyphiidae owing to the cheliceral stridulating file, the tibial autospasy, the cymbial cuspules (modified macrosetae), the retrolateral cymbial sclerite (pimoid cymbial sclerite), and the embolic process (pimoid embolic process, PEP, lost in Nanoa) (Hormiga, 1993, 2003; Hormiga et al., 2005). The placement in corroborated in several studies of Orbiculariae, among others by Griswold et al. (1998). Scharff & Coddington (1997), Coddington et al. (2004), Agnarsson (2004) and Hormiga et al. (2005).

Distribución: Western North America, Alps and Apennines, Cantabrian Mountains (northern Spain), Himalayas, Japan and adjacent islands.

Estilo de vida: Pimoids live on the underside of a fairly large sheet web, most often in wet, shady habitats, including caves.

Bibliografía:

  • Griswold, C.E., Long, C.L. & Hormiga, G. 1999. A new spider of the genus Pimoa from Gaoligong Mountains, Yunnan, China (Araneae, Araneoidea, Pimoidae). Acta Botanica Yunnanica, Suppl. 11: 91-97.

  • Hormiga, G. 1993. Implications of the phylogeny of Pimoidae for the systematics of linyphiid spiders (Araneae, Araneoidea, Linyphiidae). Mem. Qd Mus. 33: 533-542.

  • Hormiga, G. 1994a. A revision and cladistic analysis of the spider family Pimoidae (Araneoidea: Araneae). Smiths. Contr. Zool. 549: 1-104.

  • Hormiga, G. 2003. Weintrauboa, a new genus of pimoid spiders from Japan and adjacent islands, with comments on the monophyly and diagnosis of the family Pimoidae and the genus Pimoa (Araneoidea, Araneae). Zool. J. Linn. Soc. 139: 261-281.

  • Hormiga, G., Buckle, D. & Scharff, N. 2005. Nanoa, an enigmatic new genus of pimoid spiders from western North America (Pimoidae, Araneae). Zool. J. Linn. Soc. 145.

  • Hormiga, G. & Lew, S.E. 2005. Pimoidae. pp. 197-198 in Ubick, D., Paquin, P., Cushing, P.E. & Roth, V. (Eds) 2005. The Spiders of North America. An Identification Manual. American Arachnological Society.

          1 género y 1 especie presentes en la Península Ibérica:

  • Pimoa Chamberlin & Ivie, 1943

                      - P. breulli (Fage, 1931)
 
Agelena labyrinthica © Pedro Cardoso